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Name-bearing Type

• Lane and Cerqueira
• 1942:502 [pdf not available]
• Type Species: Goeldia fluviatilis (Theobald)

Classification

• Subfamily Culicinae, tribe Sabethini. Shannoniana contains only three species.

Distribution

• Species of Shannoniana occur in South and Central America.
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Phylogeny

• Shannoniana shares close affinities with other New World genera of tribe Sabethini. It was recovered as independent lineage between Johnbelkinia and Runchomyia (Ctenogoeldia) + Isostomyia in the cladistic analysis of Judd (1996) and as the sister of Isostomyia in the cladistic analyses of Harbach & Kitching (1998), Harbach & Peyton (2000) and Harbach et al. (2007a). The phylogenetic relationships of the species of Shannoniana are unknown.

Characteristics

• The adults of Shannoniana are distinguished from other members of tribe Sabethini in the New World by the following combination of characters: proboscis about as long as forefemur, broad complete or incomplete postmedian pale band on hindtibia (unique to Shannoniana), lower mesokatepisternal setae extend above lower edge of mesepimeron (distinction from Johnbelkinia, Runchomyia and Isostomyia), postprocoxal scales present and laterotergite of abdominal segment I densely and completely covered with scales (distinctions from Trichoprosopon). Shannoniana larvae are recognised by the following combination of characters: normal circular occipital foramen with a distinct collar (distinguishes Shannoniana from Isostomyia, Limatus, Phoniomyia, Sabethes, Runchomyia and most Wyeomyia), maxillary claw present and maxillary suture incomplete or absent (distinctions from Onirion, Trichoprosopon and Wyeomyia), and siphon without a line of setae on the posterior midline (distinction from Johnbelkinia). The presence of a maxillary claw distinguishes Shannoniana from all New World genera except Isostomyia, and the unusually strong articulation of the maxilla with the head capsule far ventrad of the palpus is known elsewhere only in subgenus Rachisoura of genus Tripteroides. See Sabethini.

Bionomics

• Little is known about the bionomics of Shannoniana. Larvae have been collected from bamboo and the leaf axils of aroids. Females have been captured while landing-biting on horse and humans in forest during the daytime.

Medical

• Species of Shannoniana are unlikely to be of medical or economic importance to humans.

Important References

Lane, 1953 (see next reference for included species) Zavortink, 1979 (genus as currently defined) Clark-Gil & Darsie, 1983 (Guatemala).

Included Taxa

fluviatilis (Theobald, 1903) moralesi (Dyar & Knab, 1919) schedocyclia (Dyar & Knab, 1908)

The materials presented in the Classification, Distribution, Phylogeny, Characteristics, Medical, and Important References sections, and links to subgenera, are reproduced with permission of Mosquito Taxonomic Inventory, moderated by Ralph Harbach on behalf of the contributors who retain copyright. For additional information on reuse parameters, please contact Mosquito Taxonomic Inventory. Images and maps, unless otherwise attributed, and links to the literature are provided by the WRBU.